Angolan Black & White Colobus
The Basics
The Angolan black and white colobus monkey has black hair with a white brow band, cheeks, and throat. Long haired white epaulettes stream from the shoulders.
The lower part of the tail is white as is the band on the buttocks. This subspecies, palliatus, is only found in the south eastern Kenya and Tanzania coastal forests and the Tanzanian eastern arc mountains of the East and West Usambaras, South Pares, Nguru, Nguu and Uluguru Mountains. Although the palliatus were previously found along the entire coast, deforestation and hunting in the northern parts have resulted in their restriction to isolated pockets of forests south of Mombasa. Travelling further inland, the Guereza black and white colobus occur. These are much bigger, have longer coats, have a full cape of white hair around their backs and full bushy white tail.
Here in Diani there are approximately 450 colobus monkeys identified from the annual census, carried out every October - the second highest known concentration of this subspecies in Kenya. Overall, there are an estimated 2000 remaining in Kenya. The Angolan Colobus is believed to be at risk from habitat loss and hunting. Its highly fragmented range suggests that we expect to see declining numbers in association with the further fragmentation and loss of habitat in both Kenya and Tanzania.
Colobus in Africa
Colobine monkeys (Family Cercopithecidae; Subfamily colobinae) are found in Africa and Asia. African species include the Olive, Red and Pied. The Pied colobus include the Black, Western Pied, Angola Pied, Geoffroy's Pied and the Guereza.
| Angolan Black and White Colobus | Abyssinian Black and White Colobus | Tana River Red Colobus | Zanzibar Red Colobus |
![]() |
![]() |
![]() |
![]() |
| Colobus angolensis ssp palliatus | Colobus guereza ssp occidentalis | Procolobus badius ssp tephrosceles | Procolobus badius ssp kirkii |
Note: Taxonomy of many primate species is disputed throughout the world. This is based on the World Conservation Union (ICUN) Status Survey and Conservation Primate Specialist Group.
Behaviour
As are all colobus, the Angolan colobus monkey is diurnal, they have flattened nails, pads on their buttocks, and their hind legs are longer than their fore limbs. These are typical characteristics of old world monkeys. However, the specific features of Colobines are due to their unique dietary adaptations.
Colobus eat mostly leaves (and some fruits and flowers), have no cheek pouches, are arboreal (live in the tree canopy and rarely come down to the ground) and have a light-weight bone structure and elongated limbs - making it easier to leap from branch to branch. Additionally, the Colobus have no thumbs though they retain an opposable big toe. "Colobus" in fact acquired their name from the Greek word "kolobos" meaning maimed or mutilated. The reduction of the thumb is an adaptation to arboreal living as the fingers have become aligned into a single, narrow curved arc that allows the hand to act as a flexible hook.
Their stomach is large and has three chambers, which carries specific bacteria that helps to ferment and digest leaves, similar to rumination of, for example, cows. The majority of their diet is made up of young and mature leaves - 46 species eaten but only five species make the greatest proportion of their diet. Because of the poor nutritional quality of their food, they browse intensively for many hours each day. They digest two to three kg of leaves per day (one third of their full body weight), and also eat seeds, unripe fruits and flowers. Some species of Colobus are known to eat soil, clay and charcoal which is thought to assist in the digestion of toxic leaves.
In Diani, Colobus are rarely a pest to tourists as they do not eat human food and remain in the tree canopy. Due to their dependence on forests they are a true "flagship" species in which the overall health of the forest can be gauged.
Babies and Families
Infants are born strikingly white, and then turn grey and black and then by three months of age, to the adult colouration of black and white. They are born throughout the year but a birth peak is seen in September and October. Colobine infants are known for their flamboyant colouration, which is remarkably different than the adult. This is considered an adaptation for encouraging 'aunting behaviour' where other females in the group are attracted to the newborn and spend time caring for the young. This supposedly frees up maternal time for feeding. As is known amongst the Colobines, the nutritional value to their diet is low and the stresses of rearing offspring puts enormous pressure on the female. Aunting behaviour thus counteracts the burden of nursing.
Females remain in their natal troops for life. The dominant male defends the territory and troop from predators whereas the dominant female leads the troop. Young males leave their natal troop to start bachelor groups or to travel alone until they are able to take over their own troop.
Other Primates in Diani
|
Other Wildlife in the Diani Area
|






Cercopithecus monkeys are the most common monkeys found in Africa. Five of the 20+ species found in Africa can be found in Kenya and both the vervet and sykes monkeys can be found commonly in Diani. The vervet (also known as the "African green monkey") is a grey-brown monkey with white under parts, white-fringed black face, long whitish cheek whiskers, white brow, black feet and black tip of the tail. Males posses a blue scrotum, red penis and red peri-anus and weigh around 4 to 6 kg and measure 40 to 60 cm when adult. Females are normally 2 to 5 kg and measure 30 to 50 cm. Female breasts also often have a bluish tinge to them. Infants have similar, but lighter, colouration and also have pink faces.
Sykes monkeys are also widely distributed on the Kenyan coast and are a common sight in Diani. They are approximately the same size as a Vervet, ranging from 50 to 70 cm in body length and weighing 6 to 9 kg for males and 3 to 6 kg for females.
Baboons belong to the genus Papiowhich is divided into five species - Olive, Yellow, Chacma, Guinea and Hamadryas - of which the Olive and Yellow are found in Kenya. Baboons are monkeys (although they are often wrongly referred to as apes) and are not threatened although their numbers have been declining in Africa overall in the recent years because of eradication programmes.
Galagos (or as they are more commonly known "Bush babies") are found in most forested and wooded areas in sub-Saharan Africa. They are also only found in Africa. In Diani, they frequent gardens and kitchens during the night, searching for food.
Zanj Elephant Shrew - This unique shrew (Rhynchocyon petersi) eats invertebrates in leaf litter and has been declared rare by the IUCN. Not much is known about this shrew, although they are living in the forests in Diani!
Suni - Suni are forest antelope 40cm tall and male's horns can reach 13cm. Their facial glands are enormous, especially in the male and their fur is sleek, shiny, freckled dark brown with a lighter underside. They are browsers with a varied diet of leaves, shoots and herbs and gather under feeding colobus monkeys to pick up dropped leaves and shoots of brittlewood. The rely on smell for many social activities and visible and invisible scented pathways are followed with individual and communal dung middens tending to be on the peripheries of a territory. They feed in short bursts interspersed with rests and are most active after rainstorms and between dusk and 10 pm and after 4pm. They rest during the heat of the day. Mainly pairs on territories of approximately 3 ha.
Genets and Civets - Genets and civets are to carnivores what lemurs are to the higher primates. Civets in particular are modern approximations of the carnivores common ancestral stock. These carnivores are generally solitary foragers, spanning most major habitats. Scent is their most fundamental mode of communication and all species use glandular secretions to regulate contacts and behaviour. The are almost entirely terrestrial, solitary foragers and not endangered.
African civet (Civettictis civetta) is the only civet to be found in this area. They are largely terrestrial and normally silent, although they growl very deeply if harassed. They are omnivores adapted to eating poisonous fruits, such as Strychnos, distasteful insects, millipedes and dangerous snakes. They are able to feed irregularly and even fast for two weeks at a time. They have up to four young born 60-72 days gestation born in a burrow, crevice, dense vegetation. Civet secretions are so copious and durable that they once provided the perfume trade with a valuable fixative for floral scents.
Bats - There are two types of bats in Diani - insect eaters and fruit eaters. In spite of the similarities between fruit and insectivorous bats which suggests a very ancient common ancestry, the fruit bats actual have more affinities with primates. Possibly the evolution from lemur-like gliders in rainforest. Insectivores use echo-location to pinpoint obstacles and prey. Species whose niche is under canopy utilise higher frequencies and therefore have greater precision. Those that are in open areas use lower frequencies as they require lower precision. Insect eating bats have a clawless wing, complex ears and teeth, small eyes and complex ears with irregular margins.
Frugivores (fruit-eating) bats do not use echolocation and are mediocre navigators and usually fly above the forest canopy. They have large eyes, second claw on wing, funnel-shaped ears, large tongues, blunt short teeth and a deeply ridged palate which working together crush, squeeze the fruit so that only juice and pulp are swallowed. Fibres and rind are usually spat out. They rely on a year-long supply of fruits and flowers. They can also be important as pollinators and are v. Important as seed dispersers.
Bush pigs - Bush pigs range up to 4000 m on Mt. Kilimanjaro and live in forest and woodland habitats . They are omnivorous - eating roots, tubers, bulbs, fruits, larvae, beetles, snails, amphibians and reptiles - scavenging and a group have even been seen to drive a leopard off its kill. Their home ranges are about 10km 2, nightly foraging of up to 6 km2 and are a major pest for farmers and are hunted for control and meat. Without natural predators they can become very abundant as they have short gestation periods (120 days), large litters, fast maturation rates.
Owls - Eyes can see in dim light, owls have the best hearing of all birds. Eat mice, rats, large insects. Owl can see in daylight and even on a dark night but cannot see in total darkness. Owl eyes face forward so that they can focus on their fast-moving prey and judge distances. They have large ear openings, protected by feathers to hear the noise of prey.
Baobab (Adansonia Digitata) - Truly a tree that represents Africa, this striking deciduous tree of immense girth, up to 25 meters in height. It is bare of leaves up to 9 months of the year, earning a nick-name, the upside down tree. Despite its soft pithy wood, it is one of the longest lived trees in the world. Carbon dating has shown trees 5m wide to be 1000 years old. Portuguese cannon balls from nearly four centuries ago have been found embedded in living trees that line the approach to Mombasa harbour.
Neem Tree (Azadracta indica) - Common in the coast, though it is not indigenous to Africa. It is a hardy tree which grows to 118 meters. It has a rough pale brown-grey bark, small fresh glossy-green compound leaves, small white or cream fragrant flowers and small oval greenish-yellow berries. It is widely planted at the coast for fuel, timber, shade, agroforestry in exhausted soils, and as traditional medicine for treating 40 diseases, including malaria. Also used for soap making. Colobus like to eat its leaves while its berries are a favourite for sykes and vervet monkeys.




